33
SEED TESTING INTERNATIONAL APRIL 2026
• RULES DEVELOPMENT
Validation Study for Tetrazolium Viability Test for 
Glycine max
Ignacio Aranciaga1 and Maria Belen Aranguren2
1ISTA Executive Committee Liaison Officer for ISTA Tetrazolium Committee; ignacioaranciaga@gmail.com
2ISTA Tetrazolium Committee member; maria.aranguren@corteva.com
Introduction and Objective
This study aimed to develop and statistically 
validate a tetrazolium (TZ) viability test for 
Glycine max (L.) Merr. seeds, for inclusion in 
Chapter 6 of the International Rules for Seed 
Testing (ISTA Rules). Given the economic 
importance of soybean and the need for rapid 
decision making, the objective was to establish 
a reliable, reproducible and standardised TZ 
procedure for assessing seed viability across 
laboratories.
Materials and Methods
Three soybean seed lots with different quality 
levels were selected. The study was conducted 
collaboratively by seven laboratories, including 
six ISTA accredited laboratories and one ISTA 
member laboratory. Seeds were pretreated 
by hydration for 18 h at 20 °C, between moist 
paper in sealed plastic bags. Four replicates of 
100 seeds per lot were stained for 6 h at 30 °C 
in a 1% 2,3,5-triphenyl tetrazolium chloride 
solution. Seeds were then rinsed, kept submerged 
during evaluation, manually de-coated, and 
longitudinally cut through the cotyledons and 
hypocotyl–radicle axis. Viability was assessed 
based on staining patterns, tissue turgidity, 
and location and extent of damaged or necrotic 
tissues, following the criteria of the ISTA Working 
Sheets on Tetrazolium Testing.
Results
Statistical analysis demonstrated clear 
differentiation among seed lots, with Lot 3 
showing lower viability compared to Lots 1  
and 2. Data exploration using box plots 
confirmed consistent trends across laboratories. 
All laboratory results met ISTA tolerance limits. 
Repeatability analysis showed an acceptable 
dispersion factor below 1, while reproducibility 
analysis demonstrated that interlaboratory 
variability was below the reference threshold. 
ANOVA and least squares means analysis 
confirmed statistically significant differences 
between seed lots, indicating that observed 
variability was due to seed lot quality.
Conclusions
The proposed TZ viability test for Glycine max 
meets ISTA performance criteria for repeatability 
and reproducibility. This method proved robust 
and consistent across participating laboratories, 
supporting its suitability for routine viability 
assessment. Based on the results, the proposed 
method parameters – 18-h hydration, 1% TZ 
solution, 6-h staining time and evaluation 
criteria – are recommended for inclusion in the 
ISTA Rules.
Acknowledgements
We would like to thank all the laboratories 
that participated in this validation study, as 
well as the members of the ISTA Tetrazolium 
and Statistics Committees for their valuable 
collaboration.
References
1.	ISTA Tetrazolium Committee report (1998–
2001).
2.	ISTA (2003). ISTA Working Sheets on 
Tetrazolium Testing, Vol. I. International Seed 
Testing Association, Bassersdorf, Switzerland.
3.	ISTA (2025). International Rules for Seed 
Testing, Chapter 6. International Seed Testing 
Association, Wallisellen, Switzerland.
4.	Miles, S.R. (1963). Handbook of Tolerances 
and Measures of Precision for Seed Testing. 
Proceedings of the International Seed Testing 
Association, 28(3).

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